Shaft on which a wheel turns

AXLE

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The clue "Shaft on which a wheel turns" directly describes the answer "AXLE," which is the central rod or spindle that a wheel rotates around. It’s a straightforward definition with no wordplay—just a clear reference to the mechanical part that connects wheels to a vehicle or machine. If you’ve ever seen a car lifted for repairs, the axle is the sturdy bar holding the wheels in place.